What?
Don
In a nutshell:
Modern multi-valve engines have the spark plugs in or near the center of the head, so the flame front spreads out evenly, like the Hemi. To demonstrate, take a bucket of water and drop a pebble into the middle and watch the ripples spread out from the center. Older wedges (all V8's except the Hemi) have the spark plug off to one side so that there's room for the big valves needed to get decent flow in and out of the cylinder. This puts the flame off to one side and makes it travel all the way across the cylinder to reach the mixture on the other side, which takes a long time. Same bucket, but this time drop your pebble off to one side. As with a bomb, the further you get from the ignition source, the less power the "blast" has. In this case, it's not an explosion in the cylinder (unless you have detonation, aka "knock"), it's ideally the flame front of an even burn, albeit one that happens very rapidly.
The disadvantage with the modern heads and Hemis is that the mixture doesn't swirl around inside the combustion chamber as much as it does in the Wedge head. More swirl and squish promotes more efficient burning of the mixture when the engine is running lean. To make up for it, the Hemi has to run more ignition advance (start the mixture burning sooner).
In addition, this is where modern computer-controlled electronics - both ignition and fuel injection - help; the computer can control the mixture and ignition points much more efficiently under all conditions than older mechanical systems. Mechanical systems usually have to be optimized for one condition or another. Want all out power? Emissions, fuel economy, idle quality and part-throttle driveability suffers. Want to be more usable in every day driving - gotta give up some power. That's why we now have a 425 hp 6.1L Hemi that's far more livable than the old 426 ever was and gives twice the gas mileage.
Rex, is that about right?