Well, after separating the connector, I pulled the harness out of the cable chase by the rocker panel. I inspected every inch from the front to the trunk. I was unable to find any sign of damaged insulation. The harness was actually in pretty good shape. The overbraid was dried out and brittle, and in some areas a bit crumbly. I used some flex tubing in areas where the harness passed over or around edges, or where the over braid crumbled.
I went back and looked at the connector, and low and behold, I found a layer of corrosion between the tail light supply and the two adjacent blades.
I cleaned that up, and the short was gone.
The bulkhead connector is in reasonable shape, The dash harness hasn't been hacked, except for the radio connection, and in general is in good shape with one exception. The headlight switch plug shows evidence of thermal trauma at some point before I owned the car.
So I'm not exactly convinced I've found the root cause, since the circuit I was tracing routes through the headlight switch. I find it hard to believe that the corrosive residue I cleaned up could have conducted enough current to blow the 20 amp fuse. One the other hand, depending on how much all of the bulbs take, maybe a few extra amps is enough to take the fuse over the edge.
Now that I have one short cleaned up, I'm going to go back to the front of the car to look at that portion of the harness a second time. I looked pretty closely the first time. I was monitoring resistance while I moved the harness around. But saw no change.
It's always something with an old car.